In addition to his tackles for loss, you can also add to the total his tackles for no gain and when you are done, tallying the sacks and tackles for loss and tackles for no gain it accounts for 63.5% percent of this total tackles, the highest in the NFL.
A no gain is a 'win' for the defense because even if it's 4th and 1, a no gain turns the ball over. So, it's a big play much of the time.
In terms of plays that result in no gain or a loss, excluding sacks Kawann Short is tops with 19, followed by Jurrell Casey with 17 and Damon Harrison with 16 then Donald with 14.
Here is the list of 'Percentage of tackles for no gain or loss'.